Původní popis
Meek, S. E. & S. F. Hildebrand. 1928. The marine fishes of Panama. Part III. Field Mus. Nat. Hist. Publ. Zool. Ser. v. 15 (publ. 249): xxv-xxxi + 709-1045, Pls. 72-102.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.