Původní popis
Takagi, K. 1957. Descriptions of some new gobioid fishes of Japan, with a proposition on the sensory line system as a taxonomic appliance. J. Tokyo Univ. Fish. v. 43 (no. 1): 97-126.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.